Climatic factors have the greatest overall influence on soil structure and composition.
Climate largely controls organic matter production and its rate of decay.
Climate also influences the rates of erosion and leaching.
Nevertheless, the parent material from which the soil derives does have some influence on the types of plants the soil can support.
Vegetation type has a very large impact on the kind of soil.
However, the vegetation to a large extent reflects the climate.
Soils are generally profiled in terms of their three prominent horizons: A, B and C.
The 'Aoo' layer is basically dead plant material, or mor, before it has been mixed into the soil.
Whereas the 'A' proper is the upper layer of the soil itself.
Soil layers are often visible as the gradation between the dark humus rich 'A' surface layer and the 'C' layer, which is composed mostly of parent material sans humus.
Bedrock, or other base material, composes the 'D' horizon.
Below, are some examples of common soil orders found in eastern Canada.
They are here named according to the Canadian System of Soil Classification.
The FAO Unesco system is very similar to the Canadian.
The USDA & Soil Survey's system is somewhat different.
Ontario’s boreal forests are typified by the Brunisolic and Podzolic soils.
Brunisol is a fairly rich brown soil, that corresponds to the FAO's 'Cambisol' Order.
Brown soil is commonest in the more arid west side of Ontario.
The leached podzolic soils are more prevalent in the humid east.
Luvisolic soils occur mostly in the Carolinian Forest zone.
These are the soils with clayey bases and fairly rich humus.
They have been heavily exploited as cropland.
Near the Lake Erie shore the sandy Gleysolic soils are fairly common.
These sandy loams can be excellent soils for crops.
Most of Ontario's orchards and vinyards grow on Gleysolic soils.
A few of the tropical soils are similar to soils elsewhere.
The tropical brown soils are basically similar to temperate Solonetzic, Luvisolic and Gleysolic soils.
The podzolic soils occur in the tropics also.
However, some kinds of tropical soils are quite distinctive.
One common trait of tropical soils is that the humus rots-away fairly quickly.
Basically, decomposition is more rapid in warm and wet climates.
One common feature of these soils is that they tend to range from red to chestnut-brown in colour.
These colours are due to the soils generally being very old and leached-out.
Where there is constant rainfall, both the carbonates and finer silica clays to leach away.
This makes the 'laterites' reddish, as the less-soluble sesquioxides that are left-behind are often quite reddish (Ferralsol).
If there is a long dry season, the leaching may not be too extreme.
The soils in drought-then-monsoon prone places do tend to retain a fair bit of clay.
These are the soils that crack into hexagons and crevasses during the dry seasons.
Such 'black earths' are more likely to be dark coloured, at least under the surface (Vertisol).
There are in-between soils.
Some of the tropical soils are leached and clayey. They tend to be reddish-brown.
These soils grade from chestnut-brown to quite red versions (Nitisol).
These in turn garde into the true 'red clay' soils (Acrisol).
These semi-leached red clay soils are common in the tropics and in the warmer temperate zones.
Brown soils and red soils are similar to what in the USDA system are considered to be the ‘Alfisol’ Order.
These soils are sometimes called ‘Nitosols’, but the proper FAO name is now ‘Nitisol’.
They resemble Solonetz and Luvisol, in that they have both a clayey and carbonate laden base.
They are called 'terras roxas' in parts of Brazil.
In Australia they are sometimes called 'red earths'.
Being among the best agricultural soils, they are now heavily exploited.
They are another example of how humans have changed nature.
Pristine examples of these riparian soils are becoming increasingly rare.
The first fully scientific soil classification systems were developed in the USSR.
Soil scientists (pedologists) often named the great soil groups and orders in the Russian language.
Dark prairie soil for example, is called chernozem, the Russian name for ‘black earth’.
Parkland soils sometimes have a carbonate, or even salt, hardpan layer under the humus.
This kind of soil is called solonetz, after the Russian word for ‘salty’.
Even the common word podzol is a Russian term describing the ‘ashy’ colour of the lower-layers in such soils.
Podzolic soils occur mostly under woodlands, in fairly moist climates.
In these soils, the upper ‘A’ layers are can become quite leached out.
It is largely the acidic nature of the surface humus that decelerates decay.
But the ample rain allows some leaching to occur.
Leaching accelerate the removal of clay particles and soluble minerals from the surface to the lower ‘C’ horizon.
The less soluble sesquioxides of iron and aluminium are left behind in the ‘B’ layer.
This causes the orange or reddish appearance of the ‘B’ horizon (Podzol).
The 'C' layer can accummulate the leached materials.
Sometimes even this layer becomes partly leached-out.
Mostly podzols occur under pine forests, but other kinds of sub-humid forests can also develop podzolised soil.
Something like podzolisation can occur in the rainy tropics.
In these environs, leaching can take the soluble minerals deep into the soil, even leaching the ‘C’ horizon.
Often this results in ‘red clay’ soils - which still have a fair amount of clay (Acrisol).
Taken to an extreme, laterites or latosols can form.
These laterites are very reddish in colour.
These soils even are short of clay - i.e. the finer silica particles are largely leached out.
The clays of aluminium and iron sesquioxides remain.
This makes the soil quite reddish (Ferralsol).
Such soils can harden into a rock-like crust - if they are allowed to dry-out.
These reddish soils are usually very infertile for plants.
In rainforests the tree roots are largely confined to the surface layer of humus.
Agriculture Canada. 1987.
The Canadian System of Soil Classification.
Second Edition.Canadian Government Publishing Centre.
Ottawa.
Gourou, Pierre. 1982. Terres de bonne espérance - le monde tropical. Édition Plon. Paris. pp 89-97, 192-196.
Eyre, S.R. 1968. Vegetation and Soils - a world picture. Second Edition.
Aldine Publishing Company. Chicago.
Sims, R.A., Kershaw, H.M. and Wickware, G.M.
1990.
The Autecology of Major Tree Species in the North Central Region of Ontario.
COFRDA Report 3302, NWOFTDU Technical Report 48.
Ontario Ministry of Natural Resources.
Thunder Bay.
Pender, Terry. 2003. Our Stressed-out Trees.
Ontario Arborist. International society of Arboriculture. 31(6): 10-12.
Fertiliser Excesses
Fertiliers should only be applied if they are actually required.
It is important to diagnose soil nutrient problems correctly before applying fertilisers.
When trees flush in the spring they tend to use as much of their stored carbohydrates for shoot elongation as they can afford.
If the soil is richly endowed with nutrients, a tree can act as if these nutrients surpluses are going to persist throughout the season.
When the nutrient surplus in the soil is exhausted, the tree suddenly has to revert to normal rates of growth.
Excessive fertilisation can raise the proportion of shoot growth, at the expense of the root growth.
Excessive shoot elongation can also come at the expense of the production of protective chemicals.
In other words, too much spring fertiliser can cause trees to be less drought-hardy, and less resistant to parasites and herbivores.
References
Atkins, P.W. 1995. The Periodic Kingdom.
BasicBooks. New York.
Daniel, W.D., Helms, J.A. and Baker, F.S. 1979. Principles of Silviculture.
McGraw-Hill Book Company. New York.
Emsley, John. 2001.
Nature's Building Blocks - an A-Z guide to the elements.
Oxford University Press. Oxford.
Morrison, R.T. and Boyd, R.N. 1980.
Organic Chemistry.
Allyn and Bacon Inc. Boston.
Scharenbroch, B.C. and Lloyd, J.E. 2004.
A literature review of nitrogen availability indices for use in urban landscapes.
Journal of Arboriculture. 30(4): 214-230.
Smiley, E.T. and Shirazi, A.M. 2003.
Fall fertilization and cold hardiness in landscape trees.
Journal of Arboriculture. 29(6): 342-346.
Struve, Daniel K. 2002.
A review of shade tree nitrogen fertilization research in the United States.
Journal of Arboriculture. 28(6): 252-263.
Watson, G.W. 2002. Soil replacement: long-term results.
Journal of Arboriculture. 28(5): 229-230.
Ontario’s Oak Savannah
Oak openings, oak opens, oak parkland, oak woodland and oak savannah are several names for a recurring vegetation type.
There is a strange similarity between forests on a rocky alvar in Pelee Island, the woodlands of Ontario’s Pinery Provincial Park, Toronto’s High Park, the oak openings of Missouri, Oregon’s ‘oak savanna’, and even the dry montane forests of Honduras.
These places look very much like the forest-steppes of Eurasia.
The mature forests have an oak dominated canopy, often interspersed with pine, and generally an open tallgrass under-storey.
Basically, these woodlands look like some kind of savannah.
Oak openings occur in areas that are either semi-arid, or where the soil is periodically dry.
These kinds of growing conditions occur mostly in the prairies (i.e. steppes) or on the margins thereof.
In Ontario, oak openings are commonest on old dunes, in sandy barrens, or on rocky alvars.
It is not always a lack of rain per se that explains oak openings.
Many species of oak (Quercus spp.) are well adapted to dry spells.
This is why oaks often occur where it is too dry for other kinds of trees.
It also explains how they can live on soils that are dry because of rapid drainage – even if there is a lot of rain.
Ontario's Vegetation, as it was in circa 1850.
Oak openings are a kind of the ‘parkland’ or ‘forest-steppe’ vegetation.
Oak openings grade into aspen parkland in the north.
To the south, the oak openings tend to grade into pinewoods or flatwoods.
In these parklands the tree roots spread broadly, and the root and crown competition tends to suppress the smaller groundcover plants.
In many parkland areas ground-fires were once fairly common.
These fires also helped to clear the groundcover.
In the American Midwest, burloak (Quercus macrocarpa) and chinkapin oak (Q. muhlenbergii) can dominate the oak openings.
In the eastern Great Lakes area, it is black oak (Q. velutina) that is most dominant.
There are oak openings on the margins of the palouse prairies in the Plateau Country of the western USA.
Garry oak (Q. garryana) is common in these parklands.
In the mid-latitudes of Eurasia, the pollardo oak (Q. robur) and durmast oak (Q. petraea) are very common forest-steppe species.
Tallgrass
Believe it or not, outliers of prairie once extended into southern Ontario.
They differ from the full-prairies in that Ontario is rainier.
The Great Plains can have between 100 mm and 500 mm of rain per year
Whereas, the driest corner of southern Ontario usually has at least 600 mm of rain per year.
These outlier grasslands usually have light-brown soils, and not the dark-brown soils of the Great Plains.
It is the sandy soils that seem to encourage the tallgrass - not the climate per se.
A few remnants of this tallgrass ‘prairie’ still exist in the Windsor-Essex to Pelee corridor.
A mixture of tallgrass and savannah also used to stretch between Long Point and Brantford.
Similar vegetation could be found from Toronto's lakeshore to the environs of Rice Lake.
Oak openings were most common near these patches of grassland.
Human activities, of course, have vastly reduced the oak openings as well as the tallgrass prairies.
By some estimates, less than 0.1 percent of this forest-stepe vegetation still exists in Ontario.
The best examples of tallgrass prairies occur near Lake Huron and Lake Erie.
An especially intact example of this grassland occurs in the Pinery Provincial Park, on the Huron shore.
These areas are dominated by robust grasses with wide spreading mats of rhizomes.
These tallgrasses can grow upwards of two metres tall.
The coastal sand dunes are often colonised by switch grass (Panicum virgatum).
Indian grass (Sorghastrum nutans) occurs on more established sandy areas.
Little-bluestem (Andropogon scoparius) and big-bluestem (A. gerardii) grasses are also fairly common.
These species also occur in the eastern side of the Great Plains.
Though, of course, the real prairies have a broader range of species.
Pine Barrens
In sundry parts of the world one can find forest tracts wherein most of the canopy trees are pines (Pinus).
Enigmatically, sometimes these forests seem to have little in common, except for the fact that pines are one of the dominant plants.
The hard needled pines are pre-adapted to periods of water-stress.
This is why pines are especially common on barren-lands.
They can retain their water very well during droughts.
Furthermore, being evergreen they can also take advantage of short growing seasons.
This is why pines dominate in both the high boreal forests, and in the dry parklands to the south.
Paradoxically, high alpine forests can also be pine dominated.
This is because of the rapid drainage and the short growing-hours during the day.
In the highlands of Mesoamerica and in the Greater Antilles pinewoods are common (eg. P. caribaea).
Oaks and palms often occur along with these pines.
From Alberta to Quebec, jack pine (P. banksiana) parklands occur in the dry barrens of the boreal forest.
Southern Ontario has a few white pine (P. strobus) dominated stands in the sandy barrens along the Lake Huron shoreline.
In the southern USA the ‘southern pines’ are a common sight (eg. P. palustris).
In northern Florida these pine-parklands are called the ‘flatwoods’.
Similarly, open forests dominated by Aleppo pine (P. halepensis) can occur in the drier parts of Mediterranean Europe.
These pinewoods are most common on limestone alvars or on sandy barrens.
The secret of all of these pinewoods is the pines themselves.
Pines are, in general, tolerant of water-stress and they are tolerant of short growing-periods.
Pines have some water-retaining and drought-resisting traits in common with succulents.
Thus it is not surprising that semi-arid forests and parklands tend to support pines.
At the same time, very cold locals can also become dominated by pines.
This is true in the far north, with its short growing season.
It is also true in alpine locals, where the growing-period of the day is short.
Cold air can also desiccate plants.
Both arctic and alpine plants need to be tolerant of water-stress.
Pines are paradoxical plants.
Juniper Woodlands
Pines can tolerate dry conditions, and junipers more-so.
Junipers, and to a lesser extent thujas and pines, are said to have ‘sclerophyllous’ features.
This means that these plants loose relatively little water through transpiration.
The leaves have thick cuticles, few stomata, and small surface.
The leaves may also be closely spaced, so as to cover a greater surface area on the twig.
These features may hinder photosynthesis to some extent.
Nevertheless, they also help the plants to retain their water in habitats that are often extremely arid.
Juniper woodlands tend to occur in the transitions between montane forests and the drier rain-shadows.
Such is the case on the east sides of the Santa Ana, Sierra Nevada and Cascade mountain ranges.
In the subalpine forests one may find stunted whitebark pine (P. albicaulis) or limber pine (P. flexilis), firs and other conifers.
The limiting factor is daily temperature, not rainfall.
Lower in the montane forests, between 2700 and 3200 metres altitude, there occur tall forests of ponderosa pine (P. ponderosa), Douglas fir (Pseudotsuga menziesii), and other conifers.
These montane forests have adequate rainfall (250-1000 mm/yr), at least during the winter months.
The rain-shadow vegetation varies more greatly with latitude.
East of the southern Sierra Nevada and Santa Ana mountains the montane forest grades into juniper-pinyon pine woodland.
Further down-slope, the juniper-pinyon pine woodlands grade into the Mohave Desert.
This is the desert with the diverse cacti and yuccas.
At the north end of the range there are colder winters, and somewhat more rainfall.
In the Cascade Range the rain-shadow woodlands are dominated by a mixture of junipers, pines and oaks.
These dry-woodlands grade eastward into the grass dominated Palouse prairies.
In all of these places the foothills have a sort of lower 'tree-line'.
There is an altitude below which trees give way to scrubland and/or grassland.
It is on the east side of Sierra Nevada range that juniper woodlands are most fully developed.
Western juniper (J. occidentalis) dominated forests grade into Utah juniper (J. osteosperma) woodlands.
These Utah junipers are widely spaced, due to root competition and the poor Regosolic soils.
Below about 1500 metres altitude, with less annual precipitation (100-200 mm/yr), these scrublands become nearly treeless.
Sagebrush (Artemisia tridentata) and other sclerophyllous shrubs dominate in the chaparral lands of the Great Basin.
Oddly, there are relatively few cacti in these scrublands.
In Ontario there are no chaparral lands or scrublands per se.
Nevertheless, on extremely sandy slopes, or on thin soils, junipers and white cedars can come to dominate.
In places where there were formerly sand dunes, white cedar (Thuja occidentalis) can form almost pure stands.
Thujas can tolerate rocky or sandy soils with very little humus.
On rocky alvars, or on eroded clear-cuts, red-cedars (Juniper virginiana) can be the dominant tree.
Old fields and eroded roadsides are often be colonised by stands of common juniper (J. communis).
One subspecies of common juniper (J. communis var. depressa) grows as a ground-hugging shrub.
The creeping juniper (J. horizontalis) is a prostrate shrub.
Both kinds of ground-juniper are common on rocky shorelines within the Canadian Shield.
In these cases some scrubland-like features exist on account of the poor drainage conditions.
The soil in these areas does not retain rainwater very well.
These areas are, in effect, little pockets of scrubland within the lush forests of the East.
References
Easterly, Nathan William.
1979.
Rare and Infrequent Plant Species In the Oak
Openings of Northwestern Ohio.
Ohio J. Sci. 79(2): 51-79.
Eyre, S.R. 1968. Vegetation and Soils a world picture. 2nd Edition. Aldine Publishing Company. Chicago.pp 118-121.
Munz, P.A. 1963. California Mountain Flowers. University of California Press. Berkeley.
Petrides, G.A. and Petrides, O. 1992. Western Trees. Peterson Field Guide Series. Houghton Mifflin Company. Boston.
Szeicz, J. M. and MacDonald, G. M.
1991.
Postglacial vegetation history of oak savanna in southern Ontario.
Can. J. Bot. 69(7): 1507–1519.
Tallgrass Ontario. 2009. Ontario Tallgrass. The Prairie and Savanna Association: www.TallgrassOntario.org.